I have checked the ESR on all the electrolytics, they
are all fine.
Prior to all this I had replaced C439 with a higher voltage rated
component, as recommended here:
https://trmm.net/VT100. I used a 100uF
part, rated at 100V. I had also replaced C437 as that had a high ESR
prior to seeing this problem.
Is there anything else I should check?