I'm not sure what the mystery is. It fits a Lisa,
and was used to
download software to the early prototypes of the Macintosh.
http://www.digibarn.com/collections/diagrams/early-mac-interfaces/index.html
The earliest Lisa prototypes had 50-pin edge connectors for the I/O
expansion cards. Sometime between 7/81 and 2/82 the Lisa got the
pinch-fit, 56-pin edge connector for I/O expansion cards that it
eventually shipped with. The collection of diagrams at DigiBarn (from
which the flickr image was nicked) shows this evolution pretty clearly.